About the Authors
- Philip D. Gingerich, Iyad S. Zalmout
- Museum of Paleontology and Department of Geological Sciences, University of Michigan, Ann Arbor, Michigan, United States of America
- Munir ul-Haq
- Geological Survey of Pakistan, Quetta, Pakistan
- Wighart von Koenigswald
- Steinmann-Institut für Geologie, Mineralogie und Paläontologie, Universität Bonn, Bonn, Germany
- William J. Sanders
- Museum of Paleontology, University of Michigan, Ann Arbor, Michigan, United States of America
- B. Holly Smith
- Museum of Anthropology, University of Michigan, Ann Arbor, Michigan, United States of America
The authors have declared that no competing interests exist.
Performed the experiments: ISZ. Analyzed the data: PDG MuH WvK WJS BHS. Wrote the paper: PDG MuH WvK WJS BHS ISZ. Participated in field work: PDG MuH ISZ. Contributed geological interpretations: PDG ISZ MuH. Developed chronological framework: PDG MuH ISZ. Carried out functional analysis of skeletons: PDG WJS ISZ. Interpreted life history: PDG BHS.